The LM386 amplifier's inputs are an internally biased differential pair, with common mode input range extending to the indicated ground, Vee, or actually 0.4 Volts below ground.
In order to not disturb this internal biasing, the input signal must be capacitively coupled (or perfectly ground-referenced) - a capacitor in series on the input line is needed.
Next, for a voltage gain of 200, the input signal peak to peak needs to be under 1/200 of the output voltage range of the amp. While I could not find this specified in a quick scan of the datasheet, if we assume 7 volts output voltage range with a 9 Volt supply, the input signal needs to be under 35 mV peak to peak, to avoid clipping the signal. Clipping would result in mild to severe distortion of the output - though this does not explain the complete lack of signal you find on output.
If the incoming signal is higher than this 35 mV P-P, a potentiometer as an attenuator on the input is suggested.
Once these fixes are done, please revert with results, so the answer can be added to if needed.
The solution was to physically separate the power connections for the audio components. While @Dave mentioned in the comments something about a star ground topology, the problem was that while the ground connections were all centralized, the Vcc connections were also, and shouldn't have been.
In the original design, power was supplied to the entire assembly via a small PCB with header pins for each separate component. Two rows of male header pins, one for 12V DC and one for ground. In this arrangement, all of the noise was being dumped onto the audio components each time the servo motor was actuated. Adding capacitors helped slightly, but certainly not enough.
The main two wires supplying the small power distribution PCB are about 10 cm long. I added a second pair, and created a second power distribution PCB. The original one was used for the servos and receiver, while the new one was used for audio and amplifier. The "Y" split in the power cable ultimately had the effect of keeping the power clean on the audio side.
I also removed the ground connection between the RC receiver and Arduino, though this was unnecessary. Its presence did not add noise, nor did its absence cause triggering problems.
An electrical engineer coworker advised me that it was important to have separate power supplies for motors versus digital components. While this project still only uses one power supply, the solution was to split the power as early as practical, and dedicate each to its respective purpose.
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Voltage to frequency and frequency to voltage converters are the devices I think that might be of most interest to you. The LM331 springs to mind as the V2F: -
And you can also use the LM331 to convert frequency back to voltage: -